Identifying Data 2023/24
Subject (*) Paleobiology Code 610G02043
Study programme
Grao en Bioloxía
Descriptors Cycle Period Year Type Credits
Graduate 1st four-month period
Fourth Optional 6
Language
Spanish
English
Teaching method Face-to-face
Prerequisites
Department Física e Ciencias da Terra
Coordinador
Bao Casal, Roberto
E-mail
roberto.bao@udc.es
Lecturers
Bao Casal, Roberto
Blanco Calvo, Luis Alejandro
Gonzalez Fortes, Gloria Maria
Grandal D`Anglade, Aurora
Moncunill Solé, Blanca
E-mail
roberto.bao@udc.es
alejandro.blancoc@udc.es
g.gfortes@udc.es
aurora.grandal@udc.es
blanca.moncunill@udc.es
Web http://campusvirtual.udc.es/moodle/
General description Esta materia estuda os procesos biolóxicos que operan a escala xeolóxica de tempo. Despois dunha introducción ás características principais do rexistro fósil e a súa representatividade, estúdanse aspectos relativos a análise da forma orgánica, o papel do rexistro fósil no desenvolvemento da Teoría Evolutiva moderna e a análise de procesos paleoecolóxicos e paleobioxeográficos dende una perspectiva evolutiva. No último bloque da materia preténdese obter unha visión da evolución da diversidade da vida ó longo do tempo xeolóxico no contexto dun planeta cambiante, e relacionar os coñecementos xa adquiridos, cara a interpretación da Terra como Sistema.

O enfoque da asignatura é eminentemente conceptual, deixando os aspectos mais puramente descriptivos (Paleontoloxía Sistemática) para os obradoiros e as prácticas de laboratorio.

Study programme competencies
Code Study programme competences
A1 Recoñecer distintos niveis de organización nos sistemas vivos.
A2 Identificar organismos.
A3 Recoñecer, obter, analizar e interpretar evidencias paleontológicas.
A4 Obter, manexar, conservar e observar especímenes.
A29 Impartir coñecementos de Bioloxía.
B1 Aprender a aprender.
B2 Resolver problemas de forma efectiva.

Learning aims
Learning outcomes Study programme competences
To understand the concept of deep (geologic) time A3
A29
B1
To understand the processes of fossilization and the biases of the fossil record as an indicator of ancient biospheres A2
B1
To understand how biological processes occurring at geological time scales, such as evolution or mass extinctions, cannot always be understood as simple extrapolations of processes taking place at shorter time scales A2
B1
B2
To expand our understanding of Evolutionary Theory from a multidisciplinary perspective A3
B1
B2
To know the fossil groups that make up the fossil record and their practical uses A1
A2
A3
A4
B1
B2
To identify the main bioevents in the history of the Earth, their causes and aftermath A2
A3
B1
B2
To synthesize knowledge from a long array of subjects such as Geology, Ecology, Microbiology, Biochemistry, Botany or Zoology in the framework of an ever changing Earth A2
A3
A29
B1
B2

Contents
Topic Sub-topic
SECTION-1. HISTORY AND CONCEPT OF PALEOBIOLOGY
Lesson 1. An introduction to Paleobiology 1.1 Introduction
1.2 Theoretical and methodological aspects
1.3 Divisions of Paleobiology
SECTION-2. TAPHONOMY
Lesson 2. The concept of fossil. Taphonomy 2.1 Introduction
2.2 The concept and types of fossils
2.3 Biostratinomy
2.4 Diagenesis of fossils
2.5 Ichnofossils
2.6 Time-averaging
2.7 Fossil-lagerstätten
2.8 The quality of the fossil record
SECTION-3. MORPHOLOGICAL ANALYSIS
Lesson 3. Size and Shape in Fossils 9.1 Introduction
9.2 The analysis of morphometrical variability
9.3 Types of growth
9.4 Population variability
9.5 Ecophenotypic variability
9.6 Sexual dimorphism
9.7 Taphonomical variability
Lesson 4. Ontogeny and Heterochrony 10.1 Introduction
10.2 Biogenetic and von Baer's Law
10.3 Heterochrony and its types
10.4 Heterochrony and allometry
10.5 Heterochronoclines
10.6 Dissociated heterochrony
10.7 Evolutionary consequences of heterochrony
Lesson 5. Morphodynamics and the Evolution of Form 11.1 Introduction
11.2 Constructional morphology. Phylogenetic factor. Functional factor. Fabricational factor. Other factors
11.3 Research methods in morphodynamics. Biomechanical analysis. Theoretical morphology
SECTION-4. EVOLUTIONARY PALEONTOLOGY
Lesson 6. Classification and Phylogeny 12.1 Introduction
12.2 Methods of classification. Essentialism, evolutionary, phenetic, and cladistic classification
12.3 Fossils and Phylogeny. Stratocladistics. Phylogenetic trees
Lesson 7. Speciation 13.1 Introduction
13.2 Species concepts
13.3 Modes of speciation
13.4 The problem of species concept in Paleontology
Lesson 8. Modes of evolution 14.1 Introduction
14.2 Darwinism and the Synthetic Theory of Evolution
14.3 Modes of evolution and the fossil record. Phyletic gradualism and punctuated equilibria
14.5 Evolutionary trends
14.6 Species selection
14.7 Coordinated stasis
Lesson 9. Paleobiogeography 16.1 Introduction
16.2 Dispersal biogeography
16.3 Paleogeography and paleoclimatology
16.4 Vicariance biogeography
16.5 Biogeographic patterns and extinctions
Lesson 10. Evolutionary Paleoecology 17.1 Introduction
17.2 Phanerozoic trends in global diversity. Explanatory hypotheses
17.3 Law of constant extinction. Red Queen Hypothesis and alternative explanatory hypotheses
17.4 Clade interactions
SECTION-5. BIOSTRATIGRAPHY
Lesson 11. Time and Geology 4.1 Dating methods
4.2 The geologic time scale
SECTION-6. HISTORY OF LIFE
Lesson 12. The origin and early evolution of Earth and Life 5.1 Origins of the Solar System and Earth.
5.2 Origin and evolution of the Atmosphere.
5.3 Origin of the Hidrosphere.
5.4 Origin and evolution of the continents.
5.5 The first life forms.
Lesson 13. The diversification of Life 6.1 The Ediacaran Fauna and other life forms.
6.2 The Cambrian Explosion.
6.3 Evolution of life forms during the Paleozoic.
6.4 Terrestrialization.
Lesson 14. Mass extinction events 7.1 Mass extinctions. Causes and their aftermath.
7.2 The end-Permian extinction.
7.3 The end-Cretaceous extinction.
Lesson 15. Climate and Life 8.1 Climatic evolution of the planet Earth.
8.2 Global glaciations. Methods of study.
8.3 The Snowball Earth hypothesis.
8.4 The influence of climatic change on the Quaternary faunas and floras.

Planning
Methodologies / tests Competencies Ordinary class hours Student’s personal work hours Total hours
Guest lecture / keynote speech A3 A29 B1 B2 21 63 84
Laboratory practice A1 A2 A3 A4 A29 B1 B2 14 21 35
Workshop A1 A2 A3 A4 A29 B1 B2 7 10.5 17.5
Mixed objective/subjective test A1 A2 A3 A4 A29 B1 B2 4.5 7 11.5
 
Personalized attention 2 0 2
 
(*)The information in the planning table is for guidance only and does not take into account the heterogeneity of the students.

Methodologies
Methodologies Description
Guest lecture / keynote speech Lectures will be devoted to topics related to principles and problems in Paleontology, as well as to the history of life on Earth. Students are expected to prepare their own handouts, as well as complete reading assignments from specific topics.
Laboratory practice Laboratory sessions will be devoted to the recognition of the basic morphological features of the main fossil groups, as well as to the identification of important taxa from the Iberian Peninsula. Students will be required to take their own notes and answer the lab quizzes. Unjustified changes in scheduled lab groups will not be allowed. Attendance to the lab sessions, as well as the delivery of quizzes to the lecturers, are mandatory to pass the subject.
Workshop The workshops ("clases de grupo reducido") are intended to introduce the basic concepts of taphonomy and systematics to the students by means of the direct observation of fossils. The students will prepare their own handouts and solve specific quizzes. Unjustified changes in scheduled workshop groups will not be allowed. Attendance to the workshops ("clases de grupo reducido"), as well as the delivery of quizzes to the lecturers, are mandatory to pass the subject
Mixed objective/subjective test Grading is primarily based on the idea of continuous assessment and so, the final exam IS NOT REQUIRED for those students being successful during this continuous assessment. Students failing specific parts or the whole subject are required to make the final exam for the parts they failed (see the “Assessment” section).

Personalized attention
Methodologies
Workshop
Mixed objective/subjective test
Laboratory practice
Guest lecture / keynote speech
Description
Tutoring is expected, especially for those aspects showing greater difficulty, such as quizzes solving, tests, or workshop/laboratory observations.

Part-time students not capable of attending to the workshops and/or lab sessions (i. e., those having an official academic waiver) are eligible to get an exemption of these mandatory tasks in the scheduled programme. They will be required to perform adapted specific tasks to pass the subject

Assessment
Methodologies Competencies Description Qualification
Workshop A1 A2 A3 A4 A29 B1 B2 Continuous assessment using quizzes involving multiple choice, matching, true-false questions, fill in the blank questions or short answer and essay questions. These quizzes make up 7% of the final grade 7
Mixed objective/subjective test A1 A2 A3 A4 A29 B1 B2 As stated in Step 5, grading is primarily based on the idea of continuous assessment and so, the FINAL EXAM IS NOT REQUIRED for those students being successful during this continuous assessment. For the rest of students, a final exam will be carried out for the specific parts of the subject (i. e., lectures=65%, workshops + laboratory sessions= 35%) that they failed. 0
Laboratory practice A1 A2 A3 A4 A29 B1 B2 Continuous assessment using quizzes involving multiple choice, matching, true-false questions, fill in the blank questions, short answer, essay questions and/or fossil identifications with real specimens. Quizzes make up 28% of the final grade 28
Guest lecture / keynote speech A3 A29 B1 B2 Continuous assessment will take place using in-class quizzes and participation during classes. All quizzes can involve multiple choice, matching, true-false questions, fill in the blank questions or short answer and essay questions. Quizzes make up 65% of the final grade 65
 
Assessment comments

The continuous assessment consists of:

1) Tests on the contents of lectures, making up 65% of the final grade

2) Tests on laboratory sessions + workshops, making up 35% of the final grade (tests on Systematic Paleontology, 20% + test on fossil identification “de visu”, 15%). Non-attendance to lab sessions or workshops will be penalized as follows:

  • 1 unjustified missing lab/workshop = 1 out of 10 in the Systematic Paleontology test
  • 2 unjustified missing lab/workshops = 2.5 out of 10 in the Systematic Paleontology test
  • 3 or more unjustified missing lab/workshops carries failing the entire subject (this includes the grading opportunities of January and July)

3) Besides the tests, all students are required to know the chronostratigraphic chart with 0% value in the final grade. This is therefore considered a key question to pass the subject.

Students are required to obtain a final grade (lectures, and workshops + lab sessions = 100%) of at least 5.0 out of 10 to pass this subject. However, all the activities making up the continuous assessment (lectures, and workshops + lab sessions) can be compensated among them getting a grade of at least 4.0.

The two Final Exams (grading opportunities of January and July) are only required for those students who have not passed the aforementioned continuous assessment. Students passing any of the parts of the continuous assessment (lectures, and workshops + lab sessions) are given the opportunity to keep their marks for the two Final Exams (grading opportunities of January and July), being only examined of those parts which they failed. However, all the teaching-learning process of this subject is based on the idea of being developed in the current term. This means that for successive terms the student will be required to fullfill all the assignments scheduled for those specific terms.

Under exceptional justified reasons, such as part-time learning, or students with special educational needs, specifically adapted assessments could be undertaken.

The grade “no show” will be given only to those students who have not participated in more than 20% of the assessed activities during the term.

All the aforementioned instructions also apply to part-time students.

Students from former terms, if attending the December’s advance call, will be examined under the rules of the 2022/2023 academic year (please check the corresponding syllabus).

If academic fraud is detected in any of the activities included in the continuous assessment program, the student/s involved will be subject to the current UDC regulations on this topic.


Sources of information
Basic PROTHERO, D. R. (2013). Bringing Fossils to Life. An Introduction to Paleobiology. Columbia University Press, New York
BENTON, M.J. (2020). Cowen’s History of Life. Wiley
MILSOM, C. & RIGBY, S. (2010). Fossils at a Glance. Wiley-Blackwell
DOMÈNECH, R. & MARTINELL, J (1996). Introducción a los Fósiles. Masson
BENTON, M. J. & HARPER, D. A. T. (2020). Introduction to Paleobiology and the Fossil Record. Wiiey-Blackwell
CLARKSON, E. N. K. (2001). Invertebrate Palaeontology and Evolution. Blackwell Science, Oxford
BRIGGS, D. E. G. & CROWTHER, P. R. (2003). Palaeobiology II. Blackwell Science
MARTÍNEZ-CHACÓN, M. & RIVAS, P. eds. (2009). Paleontología de Invertebrados. Sociedad Española de Paleontología
FOOTE, M. & MILLER, A.I. (2007). Principles of Paleontology. W. H. Freeman, New York
PROTHERO, D. R. (2020). The Evolving Earth. Oxford University Press
Complementary (). Digital Atlas of Ancient Life. https://www.digitalatlasofancientlife.org
(). Museo Virtual de Paleontología de la Universidad de Huelva . https://www.uhu.es/museovirtualpaleontologia/index.html
(). Paleo3D: La Colección de Prácticas Virtual del Área de Paleontología de la Universitat de València. http://paleo3d.uv.es
(). Paleobiology Database (PDBD). https://paleobiodb.org


Recommendations
Subjects that it is recommended to have taken before
Geology/610G02004
Physical Geography/610G02006
Genetics/610G02019
Population Genetics and Evolution/610G02021
Plant Systematics: Cryptogamia/610G02024
Plant Systematics: Phanerogamia/610G02025
Zoology I/610G02031
Zoology II/610G02032
Ecology I: Individuals and Ecosystems/610G02039
Ecology II: Populations and Communities/610G02040

Subjects that are recommended to be taken simultaneously
Animal Biodiversity and the Environment/610G02033

Subjects that continue the syllabus
Developmental Biology/610G02010
Functional Adaptations of Animals in the Environment/610G02037

Other comments

Students having specific inquiries or wanting to discuss class materials are always welcome during the lecturer's office hours. It is highly recommended  that they communicate any kind of problem affecting their class performance, ability to take tests or class attendances, especially in the case of foreign students.

This subject follows the Green Campus Faculty of Sciences program on sustainability (https://ciencias.udc.es/images/Facultade/Green_Campus/Declaraci%C3%B3n_Ambiental_FCiencias.pdf)



(*)The teaching guide is the document in which the URV publishes the information about all its courses. It is a public document and cannot be modified. Only in exceptional cases can it be revised by the competent agent or duly revised so that it is in line with current legislation.